The palaeoichthyological collection from the Bilovo locality in the Tver’ region (Famennian of the Main Devonian field), currently housed at the Andreapol Museum of Local History, is continued to be studied. Placoderm fishes of the family Bothriolepididae Cope predominate in the ichthyocomplex of this locality. Rare plates of large-bodied Bothriolepis sp. with a flattened armor were found at the Bilovo locality in clay deposits in addition to numerous remains of crested bothriolepidids. The material on Bothriolepis sp. is represented by three posterior medio-dorsal plates only. These plates are described, figured and compared with a similar material on Late Devonian bothriolepidids from the Main Devonian field and other regions. A brief characteristic of the Bilovo locality is given as well.
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Zhizn Zemli [Life of the Earth] 45, no 3
Zhizn Zemli [Life of the Earth] 45, no 3
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Zhizn Zemli [Life of the Earth] 45, no 3
Zhizn Zemli [Life of the Earth] 45, no 3
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10.29003/m3558.0514-7468.2023_45_3/421-432
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Moloshnikov, S.V., Linkevich, V.V.
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Keywords:
palaeontology, palaeoichthyology, placoderms, Upper Devonian, Famennian, East-European platform, Main Devonian field, Tver’ region, European Russia, museum collections.
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Moloshnikov, S.V., Linkevich, V.V., “A large-bodied non-crested antiarch (Placodermi: Bothriolepididae) from the Bilovo locality in the Tver’ region (Famennian of the Main Devonian Field)”, Zhizn Zemli [Life of the Earth] 45, no 3, 421–432 (2023) (in Russ., abstr. in Engl.). DOI: 10.29003/m3558.0514-7468.2023_45_3/421-432.
A large-bodied non-crested antiarch (Placodermi: Bothriolepididae) from the Bilovo locality in the Tver’ region (Famennian of the Main Devonian Field)
Список литературы
- Batalina, M.A., Bul’vanker, E.Z., Gekker, R.F., et al., Fauna of the Main Devonian field (Moscow–Leningrad: Academy of Sciences of USSR, 1941) (in Russian).
- Verbitsky, V.R., Verbitsky, I.V., Vasilieva, O.V., et al., State geological map of Russian Federation, Scale 1:1 000 000 (third generation), Central European part: pages O-35 – Pskov, (N-35), O-36 – Saint-Petersburg, Description, 508 p. (Saint-Petersburg: VSEGEI, 2012) (in Rus-sian).
- Vorobieva, E.I., Morphology and peculiarities of crossopterigian evolution (Moscow: Nauka, 1977) (in Russian).
- Gekker, R.F., Obruchev, D.V., Filippova, M.F., “Deposits of the Main Devonian field, V–VII”, Trudy Leningradskogo ge-ologo-gidro-geodezicheskogo tresta, 9, 82 (1935) (in Russian).
- Davydov, A.E., Linkevich, V.V., “Brachiopods of the genus Cyrtospirifer from the Bilovo formation (Famennian, Upper Devonian) in the Western Tver’ region”, Bull. Moscow Soc. Nat. Sec. Geol. 94, no 5–6, 21–32 (2019) (in Russian).
- Zhamoyda, A.I., Petrov, O.V. (eds.), Sostoyanie izuchennosti stratigrafii dokembriya i fanerozoya Rossii. Zadachi dal’neishih issledovanii. Postanovleniya Mezhvedomstvennogo stratigraficheskogo komiteta i ego postoyannih komissii, 38 (Saint-Petersburg: VSEGEI, 2008) (in Russian).
- Lebedev, O.A., Krupina, N.I., Linkevich, V.V., “First finding of a dipnoan fish (Dipnoi, Sarcopterygii) in the Famennian of the Tver’ re-gion”, Paleontologicheskii zhurnal 6, 79–90 (2019) (in Russian and English).
- Lukshevich, E.V., “Bothriolepids of the Ketleri Formation (Upper Devonian) of Latvia (Pisces, Placodermi)”, Priroda i musei, ser. paleontol. 3, 38–50 (1991) (in Russian).
- Malinovskaya, S.P., “Devonian bothriolepids (placoderm fishes) of Central Kazakhstan”, Bull. Moscow Soc. Nat. Sec. Geol. 63, no 5, 56–70 (1988) (in Russian).
- Moloshnikov, S.V., Linkevich, V.V., “New data on the Late Devonian ichthyofauna from the Tver’ region (Bilovo, Malyi Tuder river)”, In-tegrativnaya paleontologiya: perspektivi razvitiya dlya geologicheskih tselei (Saint-Petersburg, 2017) (in Russian).
- Moloshnikov, S.V., Linkevich, V.V., “Systematic position of Famennian antiarchs (Vertebrata, Placodermi) from the Tver’ region”, Izvestiya vysshikh uchebnykh zavedeniy. Geologiya i razvedka 2, 8–14 (2017) (in Russian).
- Moloshnikov, S.V., Linkevich, V.V., “Morphology and systematic position of some Famennian bothriolepidids (Placodermi, Antiarchi) from the Tver’ region”, Morphologicheskaya evolutsiya i stratigraficheskie problemi (Saint-Petersburg: VSEGEI, 2019) (in Russian).
- Moloshnikov, S.V., Linkevich, V.V., “Some notes on the variability of bothriolepidids (Placodermi, Antiarchi): an unusual specimen of Livnolepis heckeri (Lukševičs) from the Famennian of the Tver’ region”, Izvestiya visshih uchebnih zavedenii. Geologiya i razvedka 4, 5–10 (2019) (in Russian).
- Moloshnikov, S.V., Linkevich, V.V., “Late Devonian bothriolepidids (Placodermi, Antiarchi) of the Tver’ region”, Paleontol. J. 54 (2), 157–165 (2020). DOI: 10.1134/S0031030120020100.
- Moloshnikov, S.V., Linkevich, V.V., “Placoderm fishes of the family Bothriolepididae Cope (Antiarchi) from the Bilovo locality of Famennian fossils (eastern part of the Main Devonian field)”, Problemi regional’noi geologii Severnoi Evrasii. Materiali konferentsii, 72–75 (Moscow: MGRI-RGGRU, 2020) (in Russian).
- Moloshnikov, S.V., Linkevich, V.V., “Ichthyofauna (Placodermi) from the Bilovo locality of Famennian fossils in the Tver’ region: the re-sults of a study and museum exposition”, Nauka v vuzovskom musee (Moscow: MAKS Press, 2020) (in Russian).
- Obrucheva, E.D., “New bothriolepidid species from the deposits of the Zadonsk Horizon, Central Devonian field”, Problemi sovremennoi paleoihtiologii (Moscow: Nauka, 1983) (in Russian).
- Reshenie mezhvedomstvennogo regional’nogo stratigraficheskogo sovetshaniay po srednemu i verhnemu paleozoyu Russkoy platformy, s re-gional’nimi stratigraficheskimi skhemami, Leningrad, 1988: Devonakaya sistema (Leningrad, 1990) (in Russian).
- Sammet, E.Yu., “Devonian System”, Geologiya SSSR. V. 1 (Moscow: Nedra, 1971) (in Russian).
- Sammet, E.Yu., “Eastern part of the Main Devonian field”, Stratigrafiya SSSR, Devonskaya Sistema. Kniga 1 (Moscow: Nedra, 1973) (in Russian).
- Tshedukhin, A.Yu., “Upper Devonian nautiloids from the Bilovo section (Tver’ region)”, Molodye – Naukam o Zemle (Moscow: RGGRU, 2020) (in Russian).
- Downs, J.P., Daeschler, E.B., Garcia, V.E., et al., “A new large-bodied species of Bothriolepis (Antiarchi) from the Upper Devonian of Ellesmere Island, Nunavut, Canada”, J. Vert. Palaeontol. e1221833-4 (2016) DOI: 10.1080/02724634.2016.1221833.
- Gross, W., “Die Fische des Baltischen Devons”, Palaeontogr. A. 79, 97 (1933).
- Lukševičs, E., “Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the northwestern part of the East European platform”, Geodiversitas 23 (4), 489–609 (2001).
- Lukševičs, E., Beznosov, P., Stūris, V., “A new assessment of the Late Devonian antiarchan fish Bothriolepis leptocheira from South Timan (Russia) and the biotic crisis near the Frasnian–Famennian boundary”, Acta Palaeontol. Pol. 62 (1), 97–119 (2017).
- Miles, R.S., “The Old Red Sandstone antiarchs of Scotland: family Bothriolepididae”, Palaeontogr. Soc. Monogr. 130, 1–130 (1968).
- Moloshnikov, S., “Crested antiarch Bothriolepis zadonica H.D. Obrucheva from the Lower Famennian of Central European Russia”, Acta Palaeontol. Pol. 49 (1), 135–146 (2004).
- Moloshnikov, S.V., “Devonian antiarchs (Pisces, Antiarchi) from Central and Southern European Russia”, Paleontol. J. 42 (7), 691–773 (2008) DOI:10.1134/S0031030108070010.
- Moloshnikov, S.V., “Middle–Late Devonian Placoderms (Pisces: Antiarchi) from Central and Northern Asia”, Paleontol. J. 46 (10), 1097–1196 (2012). DOI: 10.1134/S0031030112100012.
- Stensiö, E., “On the Placodermi of the Upper Devonian of East Greenland II. Antiarchi: subfamily Bothriolepinae”, Palaeozool. Groenl. 2, 622 p. (1948).
References
- Batalina, M.A., Bul’vanker, E.Z., Gekker, R.F., et al., Fauna of the Main Devonian field (Moscow–Leningrad: Academy of Sciences of USSR, 1941) (in Russian).
- Verbitsky, V.R., Verbitsky, I.V., Vasilieva, O.V., et al., State geological map of Russian Federation, Scale 1:1 000 000 (third generation), Central European part: pages O-35 – Pskov, (N-35), O-36 – Saint-Petersburg, Description, 508 p. (Saint-Petersburg: VSEGEI, 2012) (in Rus-sian).
- Vorobieva, E.I., Morphology and peculiarities of crossopterigian evolution (Moscow: Nauka, 1977) (in Russian).
- Gekker, R.F., Obruchev, D.V., Filippova, M.F., “Deposits of the Main Devonian field, V–VII”, Trudy Leningradskogo ge-ologo-gidro-geodezicheskogo tresta, 9, 82 (1935) (in Russian).
- Davydov, A.E., Linkevich, V.V., “Brachiopods of the genus Cyrtospirifer from the Bilovo formation (Famennian, Upper Devonian) in the Western Tver’ region”, Bull. Moscow Soc. Nat. Sec. Geol. 94, no 5–6, 21–32 (2019) (in Russian).
- Zhamoyda, A.I., Petrov, O.V. (eds.), Sostoyanie izuchennosti stratigrafii dokembriya i fanerozoya Rossii. Zadachi dal’neishih issledovanii. Postanovleniya Mezhvedomstvennogo stratigraficheskogo komiteta i ego postoyannih komissii, 38 (Saint-Petersburg: VSEGEI, 2008) (in Russian).
- Lebedev, O.A., Krupina, N.I., Linkevich, V.V., “First finding of a dipnoan fish (Dipnoi, Sarcopterygii) in the Famennian of the Tver’ re-gion”, Paleontologicheskii zhurnal 6, 79–90 (2019) (in Russian and English).
- Lukshevich, E.V., “Bothriolepids of the Ketleri Formation (Upper Devonian) of Latvia (Pisces, Placodermi)”, Priroda i musei, ser. paleontol. 3, 38–50 (1991) (in Russian).
- Malinovskaya, S.P., “Devonian bothriolepids (placoderm fishes) of Central Kazakhstan”, Bull. Moscow Soc. Nat. Sec. Geol. 63, no 5, 56–70 (1988) (in Russian).
- Moloshnikov, S.V., Linkevich, V.V., “New data on the Late Devonian ichthyofauna from the Tver’ region (Bilovo, Malyi Tuder river)”, In-tegrativnaya paleontologiya: perspektivi razvitiya dlya geologicheskih tselei (Saint-Petersburg, 2017) (in Russian).
- Moloshnikov, S.V., Linkevich, V.V., “Systematic position of Famennian antiarchs (Vertebrata, Placodermi) from the Tver’ region”, Izvestiya vysshikh uchebnykh zavedeniy. Geologiya i razvedka 2, 8–14 (2017) (in Russian).
- Moloshnikov, S.V., Linkevich, V.V., “Morphology and systematic position of some Famennian bothriolepidids (Placodermi, Antiarchi) from the Tver’ region”, Morphologicheskaya evolutsiya i stratigraficheskie problemi (Saint-Petersburg: VSEGEI, 2019) (in Russian).
- Moloshnikov, S.V., Linkevich, V.V., “Some notes on the variability of bothriolepidids (Placodermi, Antiarchi): an unusual specimen of Livnolepis heckeri (Lukševičs) from the Famennian of the Tver’ region”, Izvestiya visshih uchebnih zavedenii. Geologiya i razvedka 4, 5–10 (2019) (in Russian).
- Moloshnikov, S.V., Linkevich, V.V., “Late Devonian bothriolepidids (Placodermi, Antiarchi) of the Tver’ region”, Paleontol. J. 54 (2), 157–165 (2020). DOI: 10.1134/S0031030120020100.
- Moloshnikov, S.V., Linkevich, V.V., “Placoderm fishes of the family Bothriolepididae Cope (Antiarchi) from the Bilovo locality of Famennian fossils (eastern part of the Main Devonian field)”, Problemi regional’noi geologii Severnoi Evrasii. Materiali konferentsii, 72–75 (Moscow: MGRI-RGGRU, 2020) (in Russian).
- Moloshnikov, S.V., Linkevich, V.V., “Ichthyofauna (Placodermi) from the Bilovo locality of Famennian fossils in the Tver’ region: the re-sults of a study and museum exposition”, Nauka v vuzovskom musee (Moscow: MAKS Press, 2020) (in Russian).
- Obrucheva, E.D., “New bothriolepidid species from the deposits of the Zadonsk Horizon, Central Devonian field”, Problemi sovremennoi paleoihtiologii (Moscow: Nauka, 1983) (in Russian).
- Reshenie mezhvedomstvennogo regional’nogo stratigraficheskogo sovetshaniay po srednemu i verhnemu paleozoyu Russkoy platformy, s re-gional’nimi stratigraficheskimi skhemami, Leningrad, 1988: Devonakaya sistema (Leningrad, 1990) (in Russian).
- Sammet, E.Yu., “Devonian System”, Geologiya SSSR. V. 1 (Moscow: Nedra, 1971) (in Russian).
- Sammet, E.Yu., “Eastern part of the Main Devonian field”, Stratigrafiya SSSR, Devonskaya Sistema. Kniga 1 (Moscow: Nedra, 1973) (in Russian).
- Tshedukhin, A.Yu., “Upper Devonian nautiloids from the Bilovo section (Tver’ region)”, Molodye – Naukam o Zemle (Moscow: RGGRU, 2020) (in Russian).
- Downs, J.P., Daeschler, E.B., Garcia, V.E., et al., “A new large-bodied species of Bothriolepis (Antiarchi) from the Upper Devonian of Ellesmere Island, Nunavut, Canada”, J. Vert. Palaeontol. e1221833-4 (2016) DOI: 10.1080/02724634.2016.1221833.
- Gross, W., “Die Fische des Baltischen Devons”, Palaeontogr. A. 79, 97 (1933).
- Lukševičs, E., “Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the northwestern part of the East European platform”, Geodiversitas 23 (4), 489–609 (2001).
- Lukševičs, E., Beznosov, P., Stūris, V., “A new assessment of the Late Devonian antiarchan fish Bothriolepis leptocheira from South Timan (Russia) and the biotic crisis near the Frasnian–Famennian boundary”, Acta Palaeontol. Pol. 62 (1), 97–119 (2017).
- Miles, R.S., “The Old Red Sandstone antiarchs of Scotland: family Bothriolepididae”, Palaeontogr. Soc. Monogr. 130, 1–130 (1968).
- Moloshnikov, S., “Crested antiarch Bothriolepis zadonica H.D. Obrucheva from the Lower Famennian of Central European Russia”, Acta Palaeontol. Pol. 49 (1), 135–146 (2004).
- Moloshnikov, S.V., “Devonian antiarchs (Pisces, Antiarchi) from Central and Southern European Russia”, Paleontol. J. 42 (7), 691–773 (2008) DOI:10.1134/S0031030108070010.
- Moloshnikov, S.V., “Middle–Late Devonian Placoderms (Pisces: Antiarchi) from Central and Northern Asia”, Paleontol. J. 46 (10), 1097–1196 (2012). DOI: 10.1134/S0031030112100012.
- Stensiö, E., “On the Placodermi of the Upper Devonian of East Greenland II. Antiarchi: subfamily Bothriolepinae”, Palaeozool. Groenl. 2, 622 p. (1948).